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  • erium by application to a mucosal surface may be advantageous in certain contexts by virtue of generating an enhanced immune response at the mucosal membrane (e.g. IgA response) in addition to a systemic response. [0064] The bacterium may be applied in a nutrient medium, i.e. medium containing a substance or substances which sustain (at least in vitro) metabolic activity in the bacterium.
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